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IUCN/SCC Otter Specialist Group Bulletin
ŠIUCN/SCC Otter Specialist Group

Volume 34(B) Special Issue (November 2017)

Citation: Camp, V.L. (2017). A Bibliography On The North American River Otter Lontra canadensis. Fourth Edition Addendum. IUCN Otter Spec. Group Bull. 34 (A): 50 - 55

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A Bibliography On The North American River Otter Lontra canadensis.  
Addendum to 2nd Edition to form 4th Edition

Victor L. Camp 1

1Bonita Springs, Florida, USA. e-mail: victorcamp300@gmail.com

Victor L. Camp.  Click for larger image

Originally published August 2013; revised February 2014 as IUCN OSG Bull. 30(A), 2013


In preparing this fourth revision new and recently discovered publications have been added to the full bibliography, but are here also presented as an addendum, allowing anyone who has created a hardcopy of the bibliography to simply print out the addendum and add it to their hardcopy.

A Bibliography on the North American River Otter (Lontra canadensis)
4th Edition
Addenda to 2nd Edition

Compiled by
Victor L. Camp

2016: Journal Articles | Web Articles | Watch List
2017 | Errata | New Titles | Watch List


Ben-David, M., L.K. Duffy, G.M. Blundell, and R.T. Bowyer. (2001). Natural exposure of coastal river otters to mercury: Relation to age, diet, and survival . Environmental Toxicology and Chemistry, 20: 1986-1992.
Bowman, J., A.G. Kidd, L.A. Nituch, C. Sadowski, and A.I. Schulte-Hostedde. (2014). Testing for Aleutian Mink Disease Virus in the river otter (Lontra canadensis) in sympatry with infected American mink (Neovison vison). Journal of Wildlife Diseases, 50 (3): 689-693.
Evans, R.D. and E.M. Addison. (1996). Spatial variation in total and methyl mercury concentrations in otter (Lutra canadensis) in central Ontario, Canada. Proceedings: 4th International Conference: Mercury as a Global Pollutant. Hamburg, 4: 406.
Polechla, P.J., Jr. and E. Carrillo-Rubio. (2009). Historic and current distributions of river otters (Lontra canadensis) and (Lontra longicaudis) in the Río Grande or Río Bravo del Norte Drainage of Colorado and New Mexico, USA and of Chihuahua, Mexico and adjacent areas. IUCN Otter Specialists Group Bulletin, 26(2): 82-96.

2016, 3rd EDITION

Baldwin, E. (2013). Activity patterns, behaviors, and population status of the North American river otter (Lontra canadensis) in a northeast coastal environment, Martha’s Vineyard, Massachusetts. M.S. Thesis, Antioch University New England. Kenne, New Hampshire.
Barding, E.E and M.J. Lacki. (2015). Occurrence of nematodes (Dracunculus spp.) in reintroduced river otters in Kentucky. Journal of the Kentucky Academy of Science, 75(1-2): 94-96
Bouley, P., M. Isadore, and T. Carroll. (2015). Return of North American river otters (Lontra canadensis) to coastal habitats of the San Francisco Bay Area, California. Northwestern Naturalist, 96(1): 1-12
Crait, J. R., A.D. McIntosh, E.C. Greiner, and M. Ben-David. (2015). The influence of changing prey availability on the prevalence of Diphyllobothrium in river otters from Yellowstone National Park. The Journal of Parasitology, 101(2): 240-243.
Day, C.C., M.D. Westover, and C. Schenck. (2015). Seasonal diet of the northern river otter (Lontra canadensis): What drives prey selection? Canadian Journal of Zoology, 93: 197-205.
Feltrop, P. D. (2015). The role of silver carp in the trophic position and diet of river otters in Illinois. M.S. Thesis, Southern Illinois University. Carbondale, Illinois.
Fimrite, P. (2015). San Francisco: Otter becomes a Bay Area sensation. Houston Chronicle, January 9.
Forman, N.S. (2015). River otter population monitoring in northeastern Pennsylvania using non-invasive genetic sampling and spatial capture-recapture models. M.S. Thesis, Pennsylvania State University, Old Main, State College, Pennsylvania.
Fretueg, G. R., T.J. Martin, C. Widga, and D.R. Ruez Jr, D. R. (2015). Summer diet characteristics of North American river otters (Lontra canadensis) in central Illinois. The American Midland Naturalist, 173(2): 294-304.
Gilbert, B. (1982). The Utterly delightful otter. Sports Illustrated, December 13, 57(25): 72
Godwin, B. L., S.E. Albeke, H.L. Bergman, A. Walters, and M. Ben-David. (2015). Density of river otters (Lontra canadensis) in relation to energy development in the Green River Basin, Wyoming. Science of The Total Environment, 532: 780-790.
Green, M. L., K. Monick, M.B. Manjerovic, J. Novakofski, and N. Mateus-Pinilla. (2015). Communication stations: cameras reveal river otter (Lontra canadensis) behavior and activity patterns at latrines. Journal of Ethology, 33: 225-234.
Kimber, K., G.V. Kollias, and E.J. Dubovi. (2000). Serological survey of selected viral agents in recently captured wild river otters (Lontra canadensis). pp. 443-448. In: Fowler, M.E. and R.E. Miller, (eds.). Zoo and Wild Animal Medicine, W.B. Saunders, Philadelphia.
Kuhn, R.A. and W. Meyer. (2010). Comparative hair structure in the Lutrinae (Carnivora: Mustelidae). Mammalia, 74: 291-303
Meronk, S.E., C.C. Day, E. Flaherty, and P.A. Zollner. (2015). Investigating the impact of invasive Asian carp on river otter diet and the native fish communities of Indiana. The Summer Undergraduate Research Fellowship (SURF) Symposium. Paper 102. Purdue University. West Lafayette, Indiana.
Moss, M.L. (2015). An ethnozooarchaelogical study of land otters and people at Kit’n’Kaboodle, (49-DIX-46). Dall Island, Alaska. BC Studies, 186: 21-55.
Mowry, R. A., T.M. Schneider, E.K. Latch, M.E. Gompper, J. Beringer, and L.S. Eggert. (2015). Genetics and the successful reintroduction of the Missouri river otter. Animal Conservation, 18(2): 196-206.
Ruez Jr., D. R. (2015). Summer diet characteristics of North American river otters (Lontra canadensis) in central Illinois. The American Midland Naturalist, 173(2): 294-304.
Russell, A. (2015). Dietary patterns of Lontra canadensis in the Lower Snohomish River Estuary, Washington. Northwest Science, 89(2): 182-187.
Savage, M. and L. Klingel. (2015). Citizen monitoring after an otter restoration (Lontra canadensis) in New Mexico, USA. IUCN Otter Specialist Group Bulletin, 32 (1): 21-24
Stokes, A.N., A.M. Ray, M.W. Buktenica, B.G. Gall, E.D. Paulson, and E.D. Brodie Jr. (2015). Otter predation on Taricha granulosa and variation in Tetrodotoxin levels with elevation. Northwestern Naturalist, 96(1): 13-21.

(as of 15 March 2016)

Publications in Preparation, Press, or Review

Fike, J.A., T.L. Serfass, and O.E. Rhodes, Jr. Assessment of genetic structure of river otter populations in Eastern North America. Cannot confirm a publication as of 15 April 2016 (VLC).
Fike, J,A., E.K. Latch, O.E. Rhodes, Jr., and T.L. Serfass. Influence of time and temperature on amplification and genotyping error of DNA derived from river otter fecal material. Cannot confirm a publication as of 15 April 2016 (VLC).
Fike, J.A., T.L. Serfass, A.S. Beheler, and O.E. Rhodes, Jr. Evaluation of preservation methods for DNA analysis of river otter scat: Probability of amplification and genotyping accuracy. Proceedings of the XI International Otter Colloquium. Cannot confirm article is in this publication (VLC).
Latch, E.K. D.G. Scognamillo, J.A. Fike, and O.E. Rhodes, Jr. Fine scale population structure of coastal and upland populations of river otters in Louisiana. Cannot confirm a publication as of 15 April 2016 (VLC).

Electronic Publications (ahead of journal publication)

Albeke, S.E., N.P. Nibbelink, and M. Ben-David, (2015). Modeling behavior of coastal river otter (Lontra canadensis) in response to prey availability in Prince William Sound, Alaska: A spatially-explicit individual-based approach. PLoS ONE 10, e0126208.doi:10.1371/journal.pone.0126208.
Timm-Davis L.L, T.J. DeWitt, and C.D. Marshal. (2015). Divergent skull morphology supports two trophic specializations in otters (Lutrinae). PLoS ONE 10 (12): e0143236.doi:10,1371/journal.pone.014323

Publications on Websites

Friis-Bastaad, E. (2015). Keeping Yukon’s river otters in the swim. Your Yukon, Yukon Research Center, Access: http://yukoncollege.yk.ca/downloads/YY_Final_otter_Jan_9_2015 
Kolba, Nikolai. (2015).
Babesia spp in North American river otter (Lontra canadensis), Beavers (Castor canadensis), muskrats (Odontra zibethicus), and mink (Neovison vison) in Pennsylvania and New Jersey. Northeast Wildlife DNA Lab.
Access: http://quantum.esu.edu/dna/research/prevalence-of-babesia-spp-infection-in-aquatic-mammals-in-pa-and-nj/


Melquist, W., S. Kempema, and E.D. Stukel. (2014?). Determination of River Otter (Lontra canadensis ) Distribution and Evaluation of Potential Sites for Population Expansion in South Dakota. The Wildlife Society, South Dakota Chapter, Annual Meeting, 24-26 February 2014.
Mowry, R., Schneider, T., Latch, E., Gompper, M., Beringer, J., and L.S. Eggert. Genetic restoration lags demographic restoration in reintroduced river otters. Animal Conservation. This article may be synonymous with the authors’ 2015 publication entitled “Genetics and the successful reintroduction of the Missouri river otter.” Animal Conservation18(2): 196-206. (above)


Beckel-Katz, A.L. (1977). Preliminary observations of the social behavior of the North American river otter. Otters: The Journal of the Otter Trust, 28-32. The author's last name should be Beckel-Kratz. The citation is corrected in this edition.
McCall, R. (1995). A novel foraging association between southern river otter (Lutra canadensis) and great egrets (Casmerodius albus). Bulletin of the British Ornithologists' Club, 116(3): 199-200. The otter species should be (Lutra longicaudus). The citation is removed from this edition.

Albeke, S.E., N.P. Nibberlink, and M. Ben-David. (2015). Modeling behavior by coastal river otter (Lontra canadensis) in response to prey availability in Prince William Sound, Alaska: A spatially- explicit individual-based approach. PLOS ONE 10(6): e0126208.doi:10.1371/journal.pone.0126208.
Bailey, J.I. (2016). Determining the impact of latitude on parturition timing in captive North American river otters: A statistical analysis of AZA studbook records. M.S., Rochester Institute of Technology. Rochester, New York.
Barocas, A., H.N. Golden, M.W. Harrington, D.B. McDonald, and M. Ben-David. (2016). Coastal
Latrine sites as social information hubs and drivers of river otter fission-fusion dynamics. Animal Behaviour, 120:103-114.
Brennan, C. (2013). Boulder confirms first North American river otter sighting in a century. Boulder DailyCamera.com, Posted: 04/02/2013.
Camp, V.L.  (2013). A bibliography on the North American river otter (Lontra canadensis). IUCN
Otter Special Group Bulletin, Special Edition 30(A): 3-41.
Camp, V.L. (2015). 2ND edition: A bibliography on the North American river otter (Lontra canadensis). IUCN Otter Special Group Bulletin, Special Edition 32(B): 3-41.
Camp, V.L. (2016). 3RD edition: A bibliography on the North American river otter (Lontra canadensis). IUCN Otter Special Group Bulletin, Special Edition 33(B): 3-47.
CWHC and Submitted by Lejeune, M. (2014). A parasite of the North American river otter: Potential cause of blindness in burbot? Canadian Wildlife Health Cooperative. Online.
Access: http://blog.healthywildlife.ca.
Dornbos, P., S. Chernyak, J. Rutkiewicz, T. Cooley, S. Strom, S. Batterman, and N. Basu. (2015). Hepatic polybrominated diphenyl ether (PBDE) levels in Wisconsin river otters (Lontra canadensis) and Michigan bald eagles (Haliaeetus leucocephalus). Journal of Great Lakes Research41: 222-227.
Dubuc, L. J. (1987). Ecology of river otters on Mount Desert Island, Maine. M.S., University of Maine, Orono, Maine.
Dubuc, L.J. (1988). OTTER: Habitat use and food habits. In: Arthur, S.M. and W.B. Krohn. (1988). An annotated bibliography of predator research in Maine, 1974-1988. Maine Agricultural Experimental Station, Technical Bulletin 132: 13-14.
Dubuc, L.J., W.B. Krohn, and R.B. Owen, Jr. (1990). Predicting Occurrence of River Otters by Habitat on Mount Desert Island, Maine. The Journal of Wildlife Management, 54(4): 594-599.
Ensor, K.E. (1991). Minnesota Pollution Control Agency 1989-1991: Contaminants in Minnesota Wildlife Study. pp 8-11. In: Addison, E.M., G.A. Fox and M. Gilbertson (eds.). Proceedings of the Expert Consultation Meeting on Mink and Otter.
Faro, J.B., R.T. Bowyer, J.W. Testa, and L.K. Duffy. (1994). River otter component of the oiled mussel-bed study. Exxon Valdez Oil Spill, State/Federal Natural Resource Damage Assessment Final Report. Restoration Study 103-3. Alaska Department of Fish and Game, 34828 Kalifornsky Beach Road, Soldotna, Alaska 99669. 10pp.
Goldman, J.G. (2016). For river otters, social life is shaped by the latrine. Scientific American, online audio recording. Online.
Access: https://www.scientificamerican.com/.../for-river-otters-social-life-is-shaped-by-the-latrine/
Hansen, H., M. Ben-David, and D.B. McDonald. (2008). Effects of genotyping protocols on success and errors in identifying individual river otters (Lontra canadensis) from their faeces. Molecular Ecology Resources, 8(2): 282-289.
Hickey, B. (2016). Yes, more otters are calling the Schuylkill River home. PhillyVoice, June 22, 2016.
Indiana Department of Natural Resources. (2017). Indiana river otter trapping: River otter trapping season. May 14, 2017. Online.
Access: https://secure.in.gov/dnr/fishwild/8449.htm
Kauffman, J. (2011). Dietary preferences, parasitic infections, and spatial dynamics of Nearctic river otters (Lontra canadensis) in Pennsylvania and New Jersey. M.S., East Stroudsburg University. East Stroudsburg, Pennsylvania. 
Klenavic, K., L. Champoux, M. O'Brien, P. Daoust, R.D. Evans, and H.E. Evans. (2008). Mercury concentrations in wild mink (Mustela vison) and river otters (Lontra canadensis) collected from eastern and Atlantic Canada: Relationship to age and parasitism. Environmental Pollution, 156(2): 359-366.
Kolba, N. (2015a). Population Genetics of the North American River Otter (Lontra canadensis). M.S.,  East Stroudsburg University. East Stroudsburg, Pennsylvania. 
Kolba, N. (2015b). Babeisa spp. in North American River Otters (Lontra canadensis), beavers (Castor canadensis), Muskrats (Ondatra zibethicus) and mink (Neovison vison) in Pennsylvania and New Jersey. A lab report on file with the Wildlife DNA Laboratory, East Stroudsburg University. East Stroudsburg, Pennsylvania.
Lander, A. (2015). Art Lander's Outdoors: Once endangered river otters now likely to be found in Kentucky for generations. Northern Kentucky Tribune, December 15. Access at www.nkentuckytribune.com.
Magoulick, D.D. (unk). Effects of otter (Lontra canadensis) predation on stream communities. Active/Completed Projects, Wildlife Management, Arkansas Cooperative Fish and Wildlife research unit. Note: this is a project outline (V. Camp)
Mullhollen, J. (2014). "CSI river otters:" Research Used DNA to Gauge Numbers. Outdoor News, March.
North Dakota Game and Fish. (2016). Game and Fish considering river otter season. Online.
Access: https://gf.nd.gov/magazine/2016/nov/otter-season
Padgett-Stewart, T.M., T.M. Wilcox, K.J. Carim, K.S. McKelvey, M.K.Young, and M.K. Schwartz. (2016). An eDNA assay for river otter detection: A tool for surveying a semi-aquatic mammal. Conservation Genetics Resources, 8(1): 5-7.
Palmer, L. (2011). Genetic Structure Analysis of North American River Otters (Lontra canadensis) in Pennsylvania and New Jersey. M.S., East Stroudsburg University. East Stroudsburg, Pennsylvania. 
Poimiroo, J. (2013). California Rambling: River Otters. Mountain Democrat, p. 4A, June 24,2013.
Scordino, J. J., P. J. Gearin, S. D. Riemer, and E. M. Iwamoto. (2016). River otter (Lontra canadensis) food habits in a Washington coast watershed: Implications for a threatened species. Northwestern Naturalist, 97(1): 36-47.
Serfass, T.L. (1984). Ecology and feeding relationships of river otter (Lutra canadensis) in northeastern Pennsylvania. M.S., East Stroudsburg University. East Stroudsburg, Pennsylvania.
Seymour, M.S., K.E. Ott, D.A. Guertin,H.N. Golden,D.B. McDonald, and M. Ben-David. (2012).
Early Holocene glacial retreat isolated populations of river otters (Lontra canadensis) along the Alaskan coast. Canadian Journal of Zoology, 90(9): 1136-1148.
Sheldon, W.G. and W.G. Toll. (1964). Feeding habits of river otter in a reservoir in central Massachusetts. Journal of Mammalogy, 45: 449-455.
Skyer, M. (2006). Food habits of a re-introduced river otter population in western New York: Annual diet, temporal and spatial variation in diet, and prey selection conclusions. M.S., Rochester Institute of Technology. Rochester, New York.
Taylor, R.T. , J.A. Wong, and T.L. Serfass. (2016). Stress levels in captured river otters (Lontra canadensis) decreased after transportation to reintroduction sites. Journal of Zoo and Wildlife Medicine, 47(4):1057-1060.
Toweill, D.E. (1974). Winter food habits of river otters in western Oregon. Journal of Wildlife Management, 38(1): 107-111.
Wilber, S.A. (2015). North American river otter (Lontra canadensis) presence and habitat analysis in Florida as compared to historical data. M.S., University of South Florida. Tampa, Florida.
Wilkie, S, (2016).  Mercury and selenium in trapped river otter (Lontra canadensis) from central Saskatchewan. An overview of a project in the Department of Biology, University of Regina. Regina, Canada.
Wilson, A., S. Davenport, Q. Yin, A. Barocas, M. Ben-David, M., and T. Goodwin. (2013). Volatile organic chemicals in secretions and excretions of Alaska river otters. In: Abstracts of papers of the American Chemical Society, Vol. 245: April. 

Lumkes R.A., S. Meronk, N. Euler, C.C. Day, and P.A. Zollner. (2016). Influence of Northern River Otter Latrine Sites upon Carnivore Activity and Species Richness. Purdue University, Indiana Academy of Science Annual Meeting.

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